The male denizens of the Toi-o-nga-rangi, the uppermost of the twelve heavens. They and the Mareikura, the female denizens, are the attendants of Io. They acts as his messengers and supervisors, and have the power to visit all the lower heavens, the earth, the spirit world, and other realms. They convey to Io reports on the conditions of things in all realms. Two important members of the Whatukura are Rehua and Ruatua.
- Best, Elsdon. (1924). The Maori. Wellington: Harry H. Tombs, pp. 66, 88, 99.